(For his prolific and imaginative studies of somatic recombination in the immune system, detailing in molecular terms the genetics of antibody diversity.)
(For his elegant genetic studies, particularly in carcinogenesis, and for developing transgenic laboratory animals for the study of cancer and other diseases.)
(For brilliantly demonstrating that the DNA responsible for antibody production is routinely reshuffled to create new genes during the lifetime of an individual.)
(For his landmark systematic clinical trials of lithium as therapy and prophylaxis for manic depressive illness, which initiated a revolution in the treatment of mental disease.)